/* 生物形态设计 - 青铜色调 */
:root {
    --bronze-primary: #8c7853;
    --bronze-secondary: #b8a17d;
    --bronze-dark: #3a3226;
    --bronze-light: #f2e9e1;
    --bronze-accent: #6b5b3d;
    --bronze-text: #4d4639;
    --bronze-organic: 12px;
    --bronze-transition: all 0.4s cubic-bezier(0.175, 0.885, 0.32, 1.275);
}

* {
    margin: 0;
    padding: 0;
    box-sizing: border-box;
    font-family: 'EB Garamond', 'Noto Serif SC', serif;
}

body {
    background-color: var(--bronze-light);
    color: var(--bronze-text);
    line-height: 1.8;
    background-image: radial-gradient(circle at 10% 20%, rgba(140, 120, 83, 0.05) 0%, rgba(140, 120, 83, 0.05) 90%);
}

a {
    text-decoration: none;
    color: var(--bronze-accent);
    transition: var(--bronze-transition);
}

a:hover {
    color: var(--bronze-primary);
    text-decoration: underline wavy;
}

.container {
    width: 90%;
    max-width: 1300px;
    margin: 0 auto;
    padding: 0 20px;
}

/* 生物形态头部设计 */
header {
    background-color: var(--bronze-dark);
    padding: 25px 0;
    position: sticky;
    top: 0;
    z-index: 1000;
    clip-path: polygon(0 0, 100% 0, 100% 90%, 50% 100%, 0 90%);
    border-bottom: 3px solid var(--bronze-primary);
}

.header-content {
    display: flex;
    justify-content: space-between;
    align-items: center;
    flex-wrap: wrap;
}

.logo {
    font-size: 2.2rem;
    font-weight: 700;
    color: var(--bronze-secondary);
    text-shadow: 2px 2px 4px rgba(0, 0, 0, 0.3);
    letter-spacing: 1px;
    position: relative;
}

.logo::after {
    content: '';
    position: absolute;
    bottom: -8px;
    left: 0;
    width: 100%;
    height: 2px;
    background: linear-gradient(90deg, var(--bronze-primary), transparent);
}

nav ul {
    display: flex;
    list-style: none;
    gap: 30px;
}

nav ul li a {
    padding: 8px 15px;
    border-radius: var(--bronze-organic);
    font-size: 1.1rem;
    color: var(--bronze-light);
    position: relative;
}

nav ul li a::before {
    content: '';
    position: absolute;
    bottom: 0;
    left: 0;
    width: 0;
    height: 2px;
    background-color: var(--bronze-secondary);
    transition: var(--bronze-transition);
}

nav ul li a:hover {
    color: var(--bronze-secondary);
    text-decoration: none;
}

nav ul li a:hover::before {
    width: 100%;
}

/* 生物形态主要内容区 */
.main-content {
    background-color: white;
    padding: 40px;
    margin: 40px 0;
    border-radius: var(--bronze-organic);
    box-shadow: 0 5px 15px rgba(0, 0, 0, 0.1);
    position: relative;
    overflow: hidden;
}

.main-content::before {
    content: '';
    position: absolute;
    top: 0;
    left: 0;
    width: 100%;
    height: 8px;
    background: linear-gradient(90deg, var(--bronze-primary), var(--bronze-accent));
}

.section-title {
    font-size: 2rem;
    margin-bottom: 30px;
    padding-bottom: 15px;
    color: var(--bronze-primary);
    position: relative;
    font-weight: 600;
}

.section-title::after {
    content: '';
    position: absolute;
    bottom: 0;
    left: 0;
    width: 100px;
    height: 3px;
    background: linear-gradient(90deg, var(--bronze-primary), transparent);
    border-radius: 3px;
}

/* 生物形态文章网格 */
.organic-grid {
    display: grid;
    grid-template-columns: repeat(auto-fill, minmax(300px, 1fr));
    gap: 30px;
}

.organic-card {
    background-color: white;
    border-radius: var(--bronze-organic);
    overflow: hidden;
    transition: var(--bronze-transition);
    box-shadow: 0 3px 10px rgba(0, 0, 0, 0.1);
    border: 1px solid rgba(140, 120, 83, 0.2);
    position: relative;
}

.organic-card:hover {
    transform: translateY(-10px) rotate(1deg);
    box-shadow: 0 15px 30px rgba(0, 0, 0, 0.15);
}

.organic-card::after {
    content: '';
    position: absolute;
    bottom: 0;
    left: 0;
    width: 100%;
    height: 5px;
    background: linear-gradient(90deg, var(--bronze-primary), var(--bronze-accent));
    transition: var(--bronze-transition);
}

.organic-card:hover::after {
    height: 8px;
}

.card-image {
    width: 100%;
    height: 200px;
    object-fit: cover;
    border-bottom: 1px solid rgba(140, 120, 83, 0.2);
}

.card-body {
    padding: 20px;
}

.card-title {
    font-size: 1.3rem;
    margin-bottom: 15px;
    color: var(--bronze-dark);
    font-weight: 600;
    display: -webkit-box;
    -webkit-line-clamp: 2;
    -webkit-box-orient: vertical;
    overflow: hidden;
}

.card-meta {
    display: flex;
    justify-content: space-between;
    font-size: 0.9rem;
    color: var(--bronze-accent);
    margin-top: 15px;
    font-style: italic;
}

/* 生物形态分类标签 */
.bio-tag {
    display: inline-block;
    padding: 3px 12px;
    background-color: var(--bronze-primary);
    color: white;
    border-radius: 15px;
    font-size: 0.8rem;
    margin-bottom: 15px;
    font-style: normal;
}

/* 文章详情页样式 */
.organic-article {
    max-width: 900px;
    margin: 0 auto;
}

.article-header {
    margin-bottom: 40px;
    text-align: center;
}

.article-title {
    font-size: 2.5rem;
    margin-bottom: 20px;
    color: var(--bronze-dark);
    line-height: 1.3;
    font-weight: 700;
}

.article-meta {
    display: flex;
    justify-content: center;
    gap: 25px;
    color: var(--bronze-accent);
    margin-bottom: 25px;
    flex-wrap: wrap;
    font-style: italic;
}

.article-image {
    width: 100%;
    max-height: 500px;
    object-fit: cover;
    border-radius: var(--bronze-organic);
    margin-bottom: 30px;
    box-shadow: 0 5px 15px rgba(0, 0, 0, 0.1);
    border: 1px solid rgba(140, 120, 83, 0.3);
}

.article-content {
    line-height: 1.9;
    font-size: 1.1rem;
}

.article-content p {
    margin-bottom: 25px;
    text-align: justify;
}

.article-content img {
    max-width: 100%;
    height: auto;
    border-radius: var(--bronze-organic);
    margin: 25px auto;
    display: block;
    box-shadow: 0 3px 10px rgba(0, 0, 0, 0.1);
}

/* 生物形态分页导航 */
.bio-pagination {
    display: flex;
    justify-content: center;
    gap: 20px;
    margin-top: 50px;
}

.bio-pagination a {
    padding: 12px 25px;
    border-radius: 30px;
    background-color: white;
    color: var(--bronze-accent);
    border: 1px solid var(--bronze-primary);
    transition: var(--bronze-transition);
    font-weight: 500;
    position: relative;
    overflow: hidden;
}

.bio-pagination a::before {
    content: '';
    position: absolute;
    top: 0;
    left: -100%;
    width: 100%;
    height: 100%;
    background: linear-gradient(90deg, transparent, rgba(140, 120, 83, 0.1), transparent);
    transition: var(--bronze-transition);
}

.bio-pagination a:hover {
    background-color: var(--bronze-primary);
    color: white;
    transform: translateY(-3px);
}

.bio-pagination a:hover::before {
    left: 100%;
}

/* 生物形态友情链接 */
.bio-links {
    background-color: white;
    padding: 30px;
    margin: 40px 0;
    border-radius: var(--bronze-organic);
    box-shadow: 0 5px 15px rgba(0, 0, 0, 0.1);
    position: relative;
}

.bio-links h3 {
    margin-bottom: 25px;
    color: var(--bronze-primary);
    font-size: 1.5rem;
}

.bio-links-container {
    display: flex;
    flex-wrap: wrap;
    gap: 15px;
}

.bio-links-container a {
    padding: 8px 20px;
    background-color: var(--bronze-light);
    border-radius: 30px;
    font-size: 0.9rem;
    color: var(--bronze-accent);
    border: 1px solid var(--bronze-primary);
    transition: var(--bronze-transition);
}

.bio-links-container a:hover {
    background-color: var(--bronze-primary);
    color: white;
    transform: translateY(-3px);
}

/* 生物形态页脚 */
footer {
    background-color: var(--bronze-dark);
    padding: 40px 0 20px;
    text-align: center;
    margin-top: 60px;
    clip-path: polygon(0 20%, 50% 0, 100% 20%, 100% 100%, 0 100%);
    position: relative;
}

footer::before {
    content: '';
    position: absolute;
    top: 0;
    left: 0;
    width: 100%;
    height: 3px;
    background: linear-gradient(90deg, var(--bronze-primary), var(--bronze-accent));
}

.copyright {
    font-size: 0.9rem;
    color: var(--bronze-secondary);
    margin-top: 30px;
}

/* 响应式设计 */
@media (max-width: 768px) {
    .header-content {
        flex-direction: column;
        gap: 20px;
    }
    
    nav ul {
        gap: 15px;
        justify-content: center;
        flex-wrap: wrap;
    }
    
    .organic-grid {
        grid-template-columns: 1fr;
    }
    
    .article-title {
        font-size: 2rem;
    }
    
    .bio-pagination {
        flex-direction: column;
        align-items: center;
    }
}